File: codeml_nssites.mlc

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Hsa_Human             AAG GTC TTT GAA AGG TGT GAG TTG GCC AGA ACT CTG AAA AGA TTG GGA ATG GAT GGC TAC AGG GGA ATC AGC CTA GCA AAC TGG ATG TGT TTG GCC AAA TGG GAG AGT GGT TAC AAC ACA CGA GCT ACA AAC TAC AAT GCT GGA GAC AGA AGC ACT GAT TAT GGG ATA TTT CAG ATC AAT AGC CGC TAC TGG TGT AAT GAT GGC AAA ACC CCA GGA GCA GTT AAT GCC TGT CAT TTA TCC TGC AGT GCT TTG CTG CAA GAT AAC ATC GCT GAT GCT GTA GCT TGT GCA AAG AGG GTT GTC CGT GAT CCA CAA GGC ATT AGA GCA TGG GTG GCA TGG AGA AAT CGT TGT CAA AAC AGA GAT GTC CGT CAG TAT GTT CAA GGT TGT GGA GTG 
Hla_gibbon            AAG GTC TTT GAA AGG TGT GAG TTG GCC AGA ACT CTG AAA AGA TTG GGA ATG GAT GGC TAC AGG GGA ATC AGC CTA GCA AAC TGG ATG TGT TTG GCC AAA TGG GAG AGT GGT TAT AAC ACA CGA GCT ACA AAC TAC AAT CCT GGA GAC AGA AGC ACT GAT TAT GGG ATA TTT CAG ATC AAT AGC CGC TAC TGG TGT AAT GAT GGC AAA ACC CCA GGA GCA GTT AAT GCC TGT CAT TTA TCC TGC AAT GCT TTG CTG CAA GAT AAC ATC GCC GAT GCT GTA GCT TGT GCA AAG AGG GTT GTC CGC GAT CCA CAA GGC ATT AGA GCA TGG GTG GCA TGG AGA AAT CGT TGT CAA AAC AGA GAT CTC CGT CAG TAT ATT CAA GGT TGT GGA GTA 
Cgu/Can_colobus       AAG ATC TTT GAA AGG TGT GAG TTG GCC AGA ACT CTG AAA AAA TTG GGA CTG GAT GGC TAC AAG GGA GTC AGC CTA GCA AAC TGG GTG TGT TTG GCC AAA TGG GAG AGT GGT TAT AAC ACA GAC GCT ACA AAC TAC AAT CCT GGA GAT GAA AGC ACT GAT TAT GGG ATA TTT CAG ATC AAT AGC CGC TAC TGG TGT AAT AAT GGC AAA ACC CCA GGA GCA GTT AAT GCC TGT CAT ATA TCC TGC AAT GCT TTG CTG CAA AAT AAC ATC GCT GAT GCT GTA GCT TGT GCA AAG AGG GTT GTC AGT GAT CCA CAA GGC ATT CGA GCA TGG GTG GCA TGG AAA AAG CAC TGT CAA AAC AGA GAT GTC AGT CAG TAT GTT GAA GGT TGT GGA GTA 
Pne_langur            AAG ATC TTT GAA AGG TGT GAG TTG GCC AGA ACT CTG AAA AAA TTG GGA CTG GAT GGC TAC AAG GGA GTC AGC CTA GCA AAC TGG GTG TGT TTG GCC AAA TGG GAG AGT GGT TAT AAC ACA GAA GCT ACA AAC TAC AAT CCT GGA GAC GAA AGC ACT GAT TAT GGG ATA TTT CAG ATC AAT AGC CGC TAC TGG TGT AAT AAT GGC AAA ACC CCA GGA GCA GTT GAT GCC TGT CAT ATA TCC TGC AGT GCT TTG CTG CAA AAC AAC ATC GCT GAT GCT GTA GCT TGT GCA AAG AGG GTT GTC AGT GAT CCA CAA GGC GTT CGA GCA TGG GTG GCA TGG AGA AAT CAC TGT CAA AAC AAA GAT GTC AGT CAG TAC GTT AAA GGT TGT GGA GTG 
Mmu_rhesus            AAG ATC TTT GAA AGG TGT GAG TTG GCC AGA ACT CTG AAA AGA TTG GGA CTG GAT GGC TAC AGG GGA ATC AGC CTA GCA AAC TGG GTG TGT TTG GCC AAA TGG GAG AGT AAT TAT AAC ACA CAA GCT ACA AAC TAC AAT CCT GGA GAC CAA AGC ACT GAT TAT GGG ATA TTT CAG ATC AAT AGC CAC TAC TGG TGT AAT AAT GGC AAA ACC CCA GGA GCA GTT AAT GCC TGT CAT ATA TCC TGC AAT GCT TTG CTG CAA GAT AAC ATC GCT GAT GCT GTA ACT TGT GCA AAG AGG GTT GTC AGT GAT CCA CAA GGC ATT AGA GCA TGG GTG GCA TGG AGA AAT CAC TGT CAA AAC AGA GAT GTC AGT CAG TAT GTT CAA GGT TGT GGA GTG 
Ssc_squirrelM         AAG GTC TTC GAA AGG TGT GAG TTG GCC AGA ACT CTG AAA AGG CTT GGA ATG GAT GGC TAC AGG GGA ATC AGC CTA GCA AAC TGG ATG TGT TTG GCC AAA TGG GAG AGT GAC TAT AAC ACA CGT GCT ACA AAC TAC AAT CCT GGA GAC CAA AGC ACT GAT TAT GGG ATA TTT CAG ATC AAT AGC CAC TAT TGG TGT AAT AAT GGC AGA ACC CCA GGA GCA GTT AAT GCC TGT CAT ATA TCC TGC AAT GCT TTG CTG CAA GAT GAC ATC ACT CAA GCT GTG GCC TGT GCA AAG AGG GTT GTC CGT GAT CCA CAA GGC ATT AGA GCA TGG GTG GCA TGG AAA GCT CAT TGT CAA AAC AGA GAT GTC AGT CAG TAT GTT CAA GGT TGT GGA GTA 
Cja_marmoset          AAG GTC TTT GAA AGG TGT GAG TTG GCC AGA ACT CTG AAA AGG TTT GGA CTG GAT GGC TAC AGG GGA ATC AGC CTA GCA AAC TGG ATG TGT TTG GCC AAA TGG GAG AGT GAT TAT AAC ACA CGT GCT ACA AAC TAC AAT CCT GGA GAC CAA AGC ACT GAT TAT GGG ATA TTT CAG ATC AAT AGC CAC TAT TGG TGT AAC AAT GGC AGA ACC CCA GGA GCA GTT AAT GCC TGT CAT ATA TCC TGC AAT GCT TTG CTG CAA GAT GAC ATC ACT GAA GCT GTG GCC TGT GCA AAG AGG GTT GTC CGC GAT CCA CAA GGC ATT AGG GCA TGG GTG GCA TGG AAA GCT CAT TGT CAA AAC AGA GAT GTC AGT CAG TAT GTT CAA GGT TGT GGA GTA 


CODONML (in paml 3.13, August 2002)    lysozymeSmall.txt   Model: One dN/dS ratio dGamma (ncatG=11) 
Codon frequencies: F3x4
Warning: Gamma model for codons.  See documentation.Site-class models

ns = 7  	ls = 130
# site patterns = 81
    2    1    1    1    2    7    2    3    3    1    2    2    2    1    1
    5    1    4    3    2    1    1    3    1    5    4    5    1    1    1
    1    2    1    3    2    1    1    1    1    1    1    1    2    2    1
    1    1    1    1    1    2    2    1    1    1    1    1    1    3    1
    1    1    1    1    1    1    1    1    1    1    1    1    1    1    1
    1    1    1    1    1    1

1       
Hsa_Human             AAG GTC TTT GAA AGG TGT GAG TTG GCC AGA ACT CTG AAA AGA TTG GGA ATG GAT GGC TAC AGG ATC AGC CTA GCA AAC TGG ATG AGT GGT TAC ACA CGA GCT AAT GCT GAC AGA TAT GGG ATA TTT CAG ATC CGC TAC AAT GAT AAA ACC CCA GTT AAT CAT TTA TCC TGC AGT CAA GAT AAC GCT GAT GTA GCT GTC CGT ATT AGA GTG AGA AAT CGT AGA GTC CGT TAT GTT CAA GGT GTG 
Hla_gibbon            ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ..T ... ... ... ... C.. ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... ... .A. ... ... ... ..C ... ... ... ... ..C ... ... ... ... ... ... ... C.. ... ... A.. ... ... ..A 
Cgu/Can_colobus       ... A.. ... ... ... ... ... ... ... ... ... ... ... .A. ... ... C.. ... ... ... .A. G.. ... ... ... ... ... G.. ... ... ..T ... GAC ... ... C.. ..T GA. ... ... ... ... ... ... ... ... ... A.. ... ... ... ... ... ... A.. ... ... .A. ... A.. ... ... ... ... ... ... A.. ... C.. ... .A. ..G .AC ... ... A.. ... ... G.. ... ..A 
Pne_langur            ... A.. ... ... ... ... ... ... ... ... ... ... ... .A. ... ... C.. ... ... ... .A. G.. ... ... ... ... ... G.. ... ... ..T ... GA. ... ... C.. ... GA. ... ... ... ... ... ... ... ... ... A.. ... ... ... ... G.. ... A.. ... ... ... ... A.C ... ... ... ... ... ... A.. G.. C.. ... ... ... .AC .A. ... A.. ..C ... A.. ... ... 
Mmu_rhesus            ... A.. ... ... ... ... ... ... ... ... ... ... ... ... ... ... C.. ... ... ... ... ... ... ... ... ... ... G.. ... AA. ..T ... .A. ... ... C.. ... CA. ... ... ... ... ... ... .A. ... ... A.. ... ... ... ... ... ... A.. ... ... .A. ... ... ... ... ... ... A.. ... A.. ... ... ... ... ... .AC ... ... A.. ... ... ... ... ... 
Ssc_squirrelM         ... ... ..C ... ... ... ... ... ... ... ... ... ... ..G C.T ... ... ... ... ... ... ... ... ... ... ... ... ... ... .AC ..T ... ..T ... ... C.. ... CA. ... ... ... ... ... ... .A. ..T ... A.. .G. ... ... ... ... ... A.. ... ... .A. ... ... G.. A.. C.A ..G ..C ... ... ... ... ... .A. GC. .A. ... ... A.. ... ... ... ... ..A 
Cja_marmoset          ... ... ... ... ... ... ... ... ... ... ... ... ... ..G ..T ... C.. ... ... ... ... ... ... ... ... ... ... ... ... .A. ..T ... ..T ... ... C.. ... CA. ... ... ... ... ... ... .A. ..T ..C A.. .G. ... ... ... ... ... A.. ... ... .A. ... ... G.. A.. ..A ..G ..C ... ..C ... ..G ... .A. GC. .A. ... ... A.. ... ... ... ... ..A 

Codon usage in sequences
--------------------------------------------------------------------------------------------------------------
Phe TTT  2  2  2  2  2  1 | Ser TCT  0  0  0  0  0  0 | Tyr TAT  2  3  3  2  3  4 | Cys TGT  7  7  7  7  7  7
    TTC  0  0  0  0  0  1 |     TCC  1  1  1  1  1  1 |     TAC  4  3  3  4  3  2 |     TGC  1  1  1  1  1  1
Leu TTA  1  1  0  0  0  0 |     TCA  0  0  0  0  0  0 | *** TAA  0  0  0  0  0  0 | *** TGA  0  0  0  0  0  0
    TTG  4  4  4  4  4  3 |     TCG  0  0  0  0  0  0 |     TAG  0  0  0  0  0  0 | Trp TGG  5  5  5  5  5  5
--------------------------------------------------------------------------------------------------------------
Leu CTT  0  0  0  0  0  1 | Pro CCT  0  1  1  1  1  1 | His CAT  1  1  1  1  1  2 | Arg CGT  3  2  0  0  0  2
    CTC  0  1  0  0  0  0 |     CCC  0  0  0  0  0  0 |     CAC  0  0  1  1  2  1 |     CGC  1  2  1  1  0  0
    CTA  1  1  1  1  1  1 |     CCA  2  2  2  2  2  2 | Gln CAA  4  4  3  3  6  6 |     CGA  1  1  1  1  0  0
    CTG  2  2  3  3  3  2 |     CCG  0  0  0  0  0  0 |     CAG  2  2  2  2  2  2 |     CGG  0  0  0  0  0  0
--------------------------------------------------------------------------------------------------------------
Ile ATT  1  2  1  0  1  1 | Thr ACT  2  2  2  2  3  3 | Asn AAT  5  6  7  5  8  6 | Ser AGT  2  1  3  4  3  2
    ATC  3  3  3  3  4  3 |     ACC  1  1  1  1  1  1 |     AAC  5  5  5  6  5  4 |     AGC  3  3  3  3  3  3
    ATA  1  1  2  2  2  2 |     ACA  2  2  2  2  2  2 | Lys AAA  3  3  5  6  3  3 | Arg AGA  6  6  2  2  5  4
Met ATG  2  2  0  0  0  2 |     ACG  0  0  0  0  0  0 |     AAG  2  2  4  3  2  2 |     AGG  3  3  2  2  3  4
--------------------------------------------------------------------------------------------------------------
Val GTT  3  2  3  4  3  3 | Ala GCT  6  4  5  5  4  4 | Asp GAT  7  7  6  6  6  5 | Gly GGT  2  2  2  2  1  1
    GTC  3  2  3  3  2  3 |     GCC  3  4  3  3  3  4 |     GAC  1  1  1  1  1  3 |     GGC  3  3  3  3  3  3
    GTA  1  2  2  1  1  1 |     GCA  5  5  5  5  5  5 | Glu GAA  1  1  3  3  1  1 |     GGA  5  5  5  5  5  5
    GTG  2  1  2  3  3  2 |     GCG  0  0  0  0  0  0 |     GAG  2  2  2  2  2  2 |     GGG  1  1  1  1  1  1
--------------------------------------------------------------------------------------------------------------

--------------------------------------------------
Phe TTT  3 | Ser TCT  0 | Tyr TAT  4 | Cys TGT  7
    TTC  0 |     TCC  1 |     TAC  2 |     TGC  1
Leu TTA  0 |     TCA  0 | *** TAA  0 | *** TGA  0
    TTG  3 |     TCG  0 |     TAG  0 | Trp TGG  5
--------------------------------------------------
Leu CTT  0 | Pro CCT  1 | His CAT  2 | Arg CGT  1
    CTC  0 |     CCC  0 |     CAC  1 |     CGC  1
    CTA  1 |     CCA  2 | Gln CAA  5 |     CGA  0
    CTG  3 |     CCG  0 |     CAG  2 |     CGG  0
--------------------------------------------------
Ile ATT  1 | Thr ACT  3 | Asn AAT  5 | Ser AGT  2
    ATC  3 |     ACC  1 |     AAC  5 |     AGC  3
    ATA  2 |     ACA  2 | Lys AAA  3 | Arg AGA  3
Met ATG  1 |     ACG  0 |     AAG  2 |     AGG  5
--------------------------------------------------
Val GTT  3 | Ala GCT  4 | Asp GAT  6 | Gly GGT  1
    GTC  3 |     GCC  4 |     GAC  2 |     GGC  3
    GTA  1 |     GCA  5 | Glu GAA  2 |     GGA  5
    GTG  2 |     GCG  0 |     GAG  2 |     GGG  1
--------------------------------------------------

Codon position x base (3x4) table for each sequence.

#1: Hsa_Human      
position  1:    T:0.20769    C:0.13077    A:0.31538    G:0.34615
position  2:    T:0.20000    C:0.16923    A:0.30000    G:0.33077
position  3:    T:0.33077    C:0.22308    A:0.25385    G:0.19231

#2: Hla_gibbon     
position  1:    T:0.20769    C:0.14615    A:0.32308    G:0.32308
position  2:    T:0.20000    C:0.16923    A:0.30769    G:0.32308
position  3:    T:0.32308    C:0.23077    A:0.26154    G:0.18462

#3: Cgu/Can_colobus
position  1:    T:0.20000    C:0.12308    A:0.32308    G:0.35385
position  2:    T:0.20000    C:0.16923    A:0.35385    G:0.27692
position  3:    T:0.33077    C:0.22308    A:0.25385    G:0.19231

#4: Pne_langur     
position  1:    T:0.20000    C:0.12308    A:0.31538    G:0.36154
position  2:    T:0.20000    C:0.16923    A:0.34615    G:0.28462
position  3:    T:0.31538    C:0.23846    A:0.25385    G:0.19231

#5: Mmu_rhesus     
position  1:    T:0.20000    C:0.13846    A:0.34615    G:0.31538
position  2:    T:0.20000    C:0.16923    A:0.34615    G:0.28462
position  3:    T:0.33077    C:0.22308    A:0.25385    G:0.19231

#6: Ssc_squirrelM  
position  1:    T:0.19231    C:0.15385    A:0.32308    G:0.33077
position  2:    T:0.20000    C:0.17692    A:0.33077    G:0.29231
position  3:    T:0.33077    C:0.23077    A:0.24615    G:0.19231

#7: Cja_marmoset   
position  1:    T:0.20000    C:0.14615    A:0.31538    G:0.33846
position  2:    T:0.20000    C:0.17692    A:0.33077    G:0.29231
position  3:    T:0.33077    C:0.23077    A:0.23846    G:0.20000

Sums of codon usage counts
------------------------------------------------------------------------------
Phe F TTT      14 | Ser S TCT       0 | Tyr Y TAT      21 | Cys C TGT      49
      TTC       1 |       TCC       7 |       TAC      21 |       TGC       7
Leu L TTA       2 |       TCA       0 | *** * TAA       0 | *** * TGA       0
      TTG      26 |       TCG       0 |       TAG       0 | Trp W TGG      35
------------------------------------------------------------------------------
Leu L CTT       1 | Pro P CCT       6 | His H CAT       9 | Arg R CGT       8
      CTC       1 |       CCC       0 |       CAC       6 |       CGC       6
      CTA       7 |       CCA      14 | Gln Q CAA      31 |       CGA       4
      CTG      18 |       CCG       0 |       CAG      14 |       CGG       0
------------------------------------------------------------------------------
Ile I ATT       7 | Thr T ACT      17 | Asn N AAT      42 | Ser S AGT      17
      ATC      22 |       ACC       7 |       AAC      35 |       AGC      21
      ATA      12 |       ACA      14 | Lys K AAA      26 | Arg R AGA      28
Met M ATG       7 |       ACG       0 |       AAG      17 |       AGG      22
------------------------------------------------------------------------------
Val V GTT      21 | Ala A GCT      32 | Asp D GAT      43 | Gly G GGT      11
      GTC      19 |       GCC      24 |       GAC      10 |       GGC      21
      GTA       9 |       GCA      35 | Glu E GAA      12 |       GGA      35
      GTG      15 |       GCG       0 |       GAG      14 |       GGG       7
------------------------------------------------------------------------------


Codon position x base (3x4) table, overall

position  1:    T:0.20110    C:0.13736    A:0.32308    G:0.33846
position  2:    T:0.20000    C:0.17143    A:0.33077    G:0.29780
position  3:    T:0.32747    C:0.22857    A:0.25165    G:0.19231

Codon frequencies under model, for use in evolver:
  0.01378574  0.00962226  0.01059374  0.00809565
  0.01181635  0.00824765  0.00908035  0.00693913
  0.02279949  0.01591374  0.00000000  0.00000000
  0.02052711  0.01432765  0.00000000  0.01205451
  0.00941649  0.00657258  0.00723616  0.00552982
  0.00807127  0.00563364  0.00620242  0.00473984
  0.01557342  0.01087004  0.01196749  0.00914546
  0.01402125  0.00978665  0.01077472  0.00823396
  0.02214758  0.01545871  0.01701945  0.01300613
  0.01898364  0.01325032  0.01458810  0.01114811
  0.03662868  0.02556633  0.02814755  0.02151013
  0.03297798  0.02301819  0.02534214  0.01936627
  0.02320222  0.01619484  0.01782990  0.01362547
  0.01988762  0.01388129  0.01528277  0.01167897
  0.03837291  0.02678377  0.02948791  0.02253443
  0.03454836  0.02411429  0.02654891  0.02028847



Nei & Gojobori 1986. dN/dS (dN, dS)
(Note: This matrix is not used in later m.l. analysis.
Use runmode = -2 for ML pairwise comparison.)

Hsa_Human           
Hla_gibbon           0.2782 (0.0133 0.0478)
Cgu/Can_colobus      1.1086 (0.0742 0.0670) 1.1055 (0.0742 0.0671)
Pne_langur           1.1979 (0.0725 0.0605) 0.9234 (0.0797 0.0863) 0.5517 (0.0267 0.0484)
Mmu_rhesus           1.8744 (0.0562 0.0300) 1.0215 (0.0561 0.0550) 1.2973 (0.0473 0.0364) 1.3970 (0.0508 0.0364)
Ssc_squirrelM        0.4701 (0.0633 0.1346) 0.4688 (0.0633 0.1349) 0.5159 (0.0775 0.1502) 0.5833 (0.0959 0.1645) 0.4544 (0.0559 0.1230)
Cja_marmoset         0.4725 (0.0634 0.1341) 0.5925 (0.0633 0.1069) 0.4702 (0.0704 0.1496) 0.5411 (0.0886 0.1638) 0.3995 (0.0490 0.1225) 0.1595 (0.0099 0.0619)


Model 0: one-ratio

TREE #  1:  ((1, 2), ((3, 4), 5), (6, 7));   MP score: 65
check convergence..
lnL(ntime: 11  np: 14):   -906.017440     +0.000000
   8..9     9..1     9..2     8..10   10..11   11..3    11..4    10..5     8..12   12..6    12..7  
  0.06798  0.02556  0.03889  0.04345  0.07637  0.04379  0.05254  0.02168  0.12266  0.04080  0.02392  4.54006  0.80663  0.50000
SEs for parameters:
  0.02557  0.01516  0.01844  0.02016  0.02668  0.01990  0.02155  0.01551  0.03392  0.01916  0.01506  0.23600  0.23600 -1.00000
Note: Branch length is defined as number of nucleotide substitutions per codon (not per neucleotide site).

tree length =   0.55765

((1: 0.02556, 2: 0.03889): 0.06798, ((3: 0.04379, 4: 0.05254): 0.07637, 5: 0.02168): 0.04345, (6: 0.04080, 7: 0.02392): 0.12266);

((Hsa_Human: 0.02556, Hla_gibbon: 0.03889): 0.06798, ((Cgu/Can_colobus: 0.04379, Pne_langur: 0.05254): 0.07637, Mmu_rhesus: 0.02168): 0.04345, (Ssc_squirrelM: 0.04080, Cja_marmoset: 0.02392): 0.12266);

Detailed output identifying parameters
kappa (ts/tv) =  4.54006

alpha (gamma) =  0.50000
r ( 1):  1.00000
f:       1.00000

dN & dS for each branch

 branch           t        S        N    dN/dS       dN       dS   S*dS   N*dN

   8..9       0.068    107.8    282.2   0.8066   0.0213   0.0263    2.8    6.0
   9..1       0.026    107.8    282.2   0.8066   0.0080   0.0099    1.1    2.3
   9..2       0.039    107.8    282.2   0.8066   0.0122   0.0151    1.6    3.4
   8..10      0.043    107.8    282.2   0.8066   0.0136   0.0168    1.8    3.8
  10..11      0.076    107.8    282.2   0.8066   0.0239   0.0296    3.2    6.7
  11..3       0.044    107.8    282.2   0.8066   0.0137   0.0170    1.8    3.9
  11..4       0.053    107.8    282.2   0.8066   0.0164   0.0204    2.2    4.6
  10..5       0.022    107.8    282.2   0.8066   0.0068   0.0084    0.9    1.9
   8..12      0.123    107.8    282.2   0.8066   0.0383   0.0475    5.1   10.8
  12..6       0.041    107.8    282.2   0.8066   0.0128   0.0158    1.7    3.6
  12..7       0.024    107.8    282.2   0.8066   0.0075   0.0093    1.0    2.1


Time used: 00:02:05


Model 1: neutral (2 categories)

TREE #  1:  ((1, 2), ((3, 4), 5), (6, 7));   MP score: 65
lnL(ntime: 11  np: 14):   -902.503869     +0.000000
   8..9     9..1     9..2     8..10   10..11   11..3    11..4    10..5     8..12   12..6    12..7  
  0.06961  0.02556  0.03938  0.04422  0.07778  0.04414  0.05228  0.02134  0.12574  0.04157  0.02367  4.29790  0.41271  0.50000
SEs for parameters:
  0.02665  0.01541  0.01888  0.02101  0.02769  0.02034  0.02188  0.01585  0.03565  0.01971  0.01530 -1.00000 -1.00000 -1.00000
Note: Branch length is defined as number of nucleotide substitutions per codon (not per neucleotide site).

tree length =   0.56528

((1: 0.02556, 2: 0.03938): 0.06961, ((3: 0.04414, 4: 0.05228): 0.07778, 5: 0.02134): 0.04422, (6: 0.04157, 7: 0.02367): 0.12574);

((Hsa_Human: 0.02556, Hla_gibbon: 0.03938): 0.06961, ((Cgu/Can_colobus: 0.04414, Pne_langur: 0.05228): 0.07778, Mmu_rhesus: 0.02134): 0.04422, (Ssc_squirrelM: 0.04157, Cja_marmoset: 0.02367): 0.12574);

Detailed output identifying parameters
kappa (ts/tv) =  4.29790


dN/dS for site classes (K=2)
p:   0.41271  0.58729
w:   0.00000  1.00000

alpha (gamma) =  0.41271
r ( 2):  0.00000  1.00000
f:       0.41271  0.58729

dN & dS for each branch

 branch           t        S        N    dN/dS       dN       dS   S*dS   N*dN

   8..9       0.070    107.1    282.9   0.5873   0.0195   0.0331    3.5    5.5
   9..1       0.026    107.1    282.9   0.5873   0.0071   0.0122    1.3    2.0
   9..2       0.039    107.1    282.9   0.5873   0.0110   0.0187    2.0    3.1
   8..10      0.044    107.1    282.9   0.5873   0.0124   0.0210    2.3    3.5
  10..11      0.078    107.1    282.9   0.5873   0.0217   0.0370    4.0    6.1
  11..3       0.044    107.1    282.9   0.5873   0.0123   0.0210    2.2    3.5
  11..4       0.052    107.1    282.9   0.5873   0.0146   0.0249    2.7    4.1
  10..5       0.021    107.1    282.9   0.5873   0.0060   0.0102    1.1    1.7
   8..12      0.126    107.1    282.9   0.5873   0.0351   0.0598    6.4    9.9
  12..6       0.042    107.1    282.9   0.5873   0.0116   0.0198    2.1    3.3
  12..7       0.024    107.1    282.9   0.5873   0.0066   0.0113    1.2    1.9


Time used: 00:05:26


Model 2: selection (3 categories)

TREE #  1:  ((1, 2), ((3, 4), 5), (6, 7));   MP score: 65
check convergence..
lnL(ntime: 11  np: 16):   -900.076500     +0.000000
   8..9     9..1     9..2     8..10   10..11   11..3    11..4    10..5     8..12   12..6    12..7  
  0.07903  0.02598  0.04137  0.04535  0.08550  0.04486  0.05417  0.01935  0.13948  0.04319  0.02492  5.12250  0.38160  0.54916  6.17292  0.41271
SEs for parameters:
  0.03226  0.01618  0.02021  0.02390  0.03229  0.02158  0.02363  0.01746  0.04358  0.02088  0.01602  1.50414  0.14016  0.16630  4.30113 -1.00000
Note: Branch length is defined as number of nucleotide substitutions per codon (not per neucleotide site).

tree length =   0.60321

((1: 0.02598, 2: 0.04137): 0.07903, ((3: 0.04486, 4: 0.05417): 0.08550, 5: 0.01935): 0.04535, (6: 0.04319, 7: 0.02492): 0.13948);

((Hsa_Human: 0.02598, Hla_gibbon: 0.04137): 0.07903, ((Cgu/Can_colobus: 0.04486, Pne_langur: 0.05417): 0.08550, Mmu_rhesus: 0.01935): 0.04535, (Ssc_squirrelM: 0.04319, Cja_marmoset: 0.02492): 0.13948);

Detailed output identifying parameters
kappa (ts/tv) =  5.12250


dN/dS for site classes (K=3)
p:   0.38160  0.54916  0.06924
w:   0.00000  1.00000  6.17292

alpha (gamma) =  0.38160
r ( 3):  0.00000  1.00000  6.17292
f:       0.38160  0.54916  0.06924

dN & dS for each branch

 branch           t        S        N    dN/dS       dN       dS   S*dS   N*dN

   8..9       0.079    109.3    280.7   0.9766   0.0262   0.0268    2.9    7.3
   9..1       0.026    109.3    280.7   0.9766   0.0086   0.0088    1.0    2.4
   9..2       0.041    109.3    280.7   0.9766   0.0137   0.0140    1.5    3.8
   8..10      0.045    109.3    280.7   0.9766   0.0150   0.0154    1.7    4.2
  10..11      0.086    109.3    280.7   0.9766   0.0283   0.0290    3.2    7.9
  11..3       0.045    109.3    280.7   0.9766   0.0149   0.0152    1.7    4.2
  11..4       0.054    109.3    280.7   0.9766   0.0179   0.0184    2.0    5.0
  10..5       0.019    109.3    280.7   0.9766   0.0064   0.0066    0.7    1.8
   8..12      0.139    109.3    280.7   0.9766   0.0462   0.0473    5.2   13.0
  12..6       0.043    109.3    280.7   0.9766   0.0143   0.0146    1.6    4.0
  12..7       0.025    109.3    280.7   0.9766   0.0083   0.0085    0.9    2.3



Positively selected sites Prob(w>1):

    15 L 0.6588
    17 M 0.6601
    37 G 0.7155
    41 R 0.9282
    50 R 0.8292
    62 R 0.5229
   114 N 0.6491
   126 Q 0.5226


Time used: 00:13:43