File: baseml_SE.out

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seed used = 1

BASEML (in paml version 4.3, August 2009)  ../Tests/PAML/alignment.phylip  REV dGamma (ncatG=5)
Frequencies..
                                    T      C      A      G
Homo_sapie                     0.2027 0.1622 0.3694 0.2658
Pan_troglo                     0.1982 0.1667 0.3694 0.2658
Gorilla_go                     0.2072 0.1577 0.3694 0.2658
Pongo_pygm                     0.2027 0.1622 0.3694 0.2658
Macaca_mul                     0.1937 0.1667 0.3739 0.2658

Homogeneity statistic: X2 = 0.00092 G = 0.00092 

Average                        0.200901 0.163063 0.370270 0.265766

# constant sites:    218 (98.20%)
ln Lmax (unconstrained) = -316.049385

Distances: TN93 (kappa)  (alpha set at 0.50)
This matrix is not used in later m.l. analysis.

Homo_sapie       
Pan_troglo         0.0047(999.0000)
Gorilla_go         0.0047(999.0000)  0.0097(999.0000)
Pongo_pygm         0.0000(999.0000)  0.0047(999.0000)  0.0047(999.0000)
Macaca_mul         0.0093( 3.6925)  0.0145( 7.6730)  0.0145( 7.7118)  0.0093( 3.6925)

TREE #  1:  (((1, 2), 3), 4, 5);  MP score: 4.00
lnL(ntime:  7  np: 13):   -319.774326    +0.000000
   6..7     7..8     8..1     8..2     7..3     6..4     6..5  
 0.000004 0.000004 0.000004 0.004533 0.004529 0.000004 0.009174 999.000000 160.593137 0.000010 0.000010 0.000010 200.111998
SEs for parameters:
 -1.00000 -1.00000 -1.00000 801727.63247 730462.67590 -1.00000 1610106.73811 -1.00000 -1.00000 3749008252.60285 16592006264.44326 4678796535.94680 212206538264286.93750

tree length =   0.01825

(((Homo_sapie, Pan_troglo), Gorilla_go), Pongo_pygm, Macaca_mul);

(((Homo_sapie: 0.000004, Pan_troglo: 0.004533): 0.000004, Gorilla_go: 0.004529): 0.000004, Pongo_pygm: 0.000004, Macaca_mul: 0.009174);

Detailed output identifying parameters

Parameters  in the rate matrix (REV) (Yang 1994 J Mol Evol 39:105-111):


Rate parameters:   999.00000 160.59314  0.00001  0.00001  0.00001
Base frequencies:   0.20090  0.16306  0.37027  0.26577
Rate matrix Q, Average Ts/Tv =   2.7478
   -2.483319    1.819246    0.664073    0.000000
    2.241391   -2.241391    0.000000    0.000000
    0.360312    0.000000   -0.363280    0.002968
    0.000000    0.000000    0.004135   -0.004135

alpha (gamma, K=5) = 200.11200
rate:   0.90307  0.96129  0.99837  1.03642  1.10086
freq:   0.20000  0.20000  0.20000  0.20000  0.20000