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\name{plot.phylo}
\alias{plot.phylo}
\alias{plot.multiPhylo}
\title{Plot Phylogenies}
\description{
These functions plot phylogenetic trees.
}
\usage{
\method{plot}{phylo}(x, type = "phylogram", use.edge.length = TRUE,
node.pos = NULL, show.tip.label = TRUE,
show.node.label = FALSE, edge.color = NULL, edge.width
= NULL, edge.lty = NULL, node.color = NULL, node.width
= NULL, node.lty = NULL, font = 3, cex = par("cex"),
adj = NULL, srt = 0, no.margin = FALSE, root.edge =
FALSE, label.offset = 0, underscore = FALSE, x.lim =
NULL, y.lim = NULL, direction = "rightwards", lab4ut =
NULL, tip.color = par("col"), plot = TRUE, rotate.tree
= 0, open.angle = 0, node.depth = 1, align.tip.label =
FALSE, ...)
\method{plot}{multiPhylo}(x, layout = 1, ...)
}
\arguments{
\item{x}{an object of class \code{"phylo"} or of class
\code{"multiPhylo"}.}
\item{type}{a character string specifying the type of phylogeny to be
drawn; it must be one of "phylogram" (the default), "cladogram",
"fan", "unrooted", "radial", "tidy", or any unambiguous abbreviation
of these.}
\item{use.edge.length}{a logical indicating whether to use the edge
lengths of the phylogeny to draw the branches (the default) or not
(if \code{FALSE}). This option has no effect if the object of class
\code{"phylo"} has no `edge.length' element.}
\item{node.pos}{a numeric taking the value 1 or 2 which specifies the
vertical position of the nodes with respect to their descendants. If
\code{NULL} (the default), then the value is determined in relation
to `type' and `use.edge.length' (see details).}
\item{show.tip.label}{a logical indicating whether to show the tip
labels on the phylogeny (defaults to \code{TRUE}, i.e. the labels
are shown).}
\item{show.node.label}{a logical indicating whether to show the node
labels on the phylogeny (defaults to \code{FALSE}, i.e. the labels
are not shown).}
\item{edge.color}{a vector of mode character giving the colours used
to draw the branches of the plotted phylogeny. These are taken to be
in the same order than the component \code{edge} of \code{phy}. If
fewer colours are given than the length of \code{edge}, then the
colours are recycled.}
\item{edge.width}{a numeric vector giving the width of the branches of
the plotted phylogeny. These are taken to be in the same order than
the component \code{edge} of \code{phy}. If fewer widths are given
than the length of \code{edge}, then these are recycled.}
\item{edge.lty}{same as the previous argument but for line types;
1: plain, 2: dashed, 3: dotted, 4: dotdash, 5: longdash, 6: twodash.}
\item{node.color}{a vector of mode character giving the colours used
to draw the perpendicular lines associated with each node of the
plotted phylogeny. These are taken to be
in the same order than the component \code{node} of \code{phy}. If
fewer colours are given than the length of \code{node}, then the
colours are recycled.}
\item{node.width}{as the previous argument, but for line widths.}
\item{node.lty}{as the previous argument, but for line types;
1: plain, 2: dashed, 3: dotted, 4: dotdash, 5: longdash, 6: twodash.}
\item{font}{an integer specifying the type of font for the labels: 1
(plain text), 2 (bold), 3 (italic, the default), or 4 (bold
italic).}
\item{cex}{a numeric value giving the factor scaling of the tip and
node labels (Character EXpansion). The default is to take the
current value from the graphical parameters.}
\item{adj}{a numeric specifying the justification of the text strings
of the labels: 0 (left-justification), 0.5 (centering), or 1
(right-justification). This option has no effect if \code{type =
"unrooted"}. If \code{NULL} (the default) the value is set with
respect of \code{direction} (see details).}
\item{srt}{a numeric giving how much the labels are rotated in degrees
(negative values are allowed resulting in clock-like rotation); the
value has an effect respectively to the value of
\code{direction} (see Examples). This option has no effect if
\code{type = "unrooted"}.}
\item{no.margin}{a logical. If \code{TRUE}, the margins are set to
zero and the plot uses all the space of the device (note that this
was the behaviour of \code{plot.phylo} up to version 0.2-1 of `ape'
with no way to modify it by the user, at least easily).}
\item{root.edge}{a logical indicating whether to draw the root edge
(defaults to FALSE); this has no effect if `use.edge.length = FALSE'
or if `type = "unrooted"'.}
\item{label.offset}{a numeric giving the space between the nodes and
the tips of the phylogeny and their corresponding labels. This
option has no effect if \code{type = "unrooted"}.}
\item{underscore}{a logical specifying whether the underscores in tip
labels should be written as spaces (the default) or left as are (if
\code{TRUE}).}
\item{x.lim}{a numeric vector of length one or two giving the limit(s)
of the x-axis. If \code{NULL}, this is computed with respect to
various parameters such as the string lengths of the labels and the
branch lengths. If a single value is given, this is taken as the
upper limit.}
\item{y.lim}{same than above for the y-axis.}
\item{direction}{a character string specifying the direction of the
tree. Four values are possible: "rightwards" (the default),
"leftwards", "upwards", and "downwards".}
\item{lab4ut}{(= labels for unrooted trees) a character string
specifying the display of tip labels for unrooted trees (can be
abbreviated): either \code{"horizontal"} where all labels are
horizontal (the default if \code{type = "u"}), or \code{"axial"}
where the labels are displayed in the axis of the corresponding
terminal branches. This option has an effect if \code{type = "u"},
\code{"f"}, or \code{"r"}.}
\item{tip.color}{the colours used for the tip labels, eventually
recycled (see examples).}
\item{plot}{a logical controlling whether to draw the tree. If
\code{FALSE}, the graphical device is set as if the tree was
plotted, and the coordinates are saved as well.}
\item{rotate.tree}{for "fan", "unrooted", or "radial" trees: the
rotation of the whole tree in degrees (negative values are
accepted).}
\item{open.angle}{if \code{type = "f"} or \code{"r"}, the angle in
degrees left blank. Use a non-zero value if you want to call
\code{\link{axisPhylo}} after the tree is plotted.}
\item{node.depth}{an integer value (1 or 2) used if branch lengths are
not used to plot the tree; 1: the node depths are proportional to
the number of tips descending from each node (the default and was the
only possibility previously), 2: they are evenly spaced.}
\item{align.tip.label}{a logical value or an integer. If \code{TRUE},
the tips are aligned and dotted lines are drawn between the tips of
the tree and the labels. If an integer, the tips are aligned and
this gives the type of the lines (\code{lty}).}
\item{layout}{the number of trees to be plotted simultaneously.}
\item{\dots}{further arguments to be passed to \code{plot} or to
\code{plot.phylo}.}
}
\details{
If \code{x} is a list of trees (i.e., an object of class
\code{"multiPhylo"}), then any further argument may be passed with
\code{...} and could be any one of those listed above for a single
tree.
The font format of the labels of the nodes and the tips is the same.
If \code{no.margin = TRUE}, the margins are set to zero and are not
restored after plotting the tree, so that the user can access the
coordinates system of the plot.
The option `node.pos' allows the user to alter the vertical position
(i.e., ordinates) of the nodes. If \code{node.pos = 1}, then the
ordinate of a node is the mean of the ordinates of its direct
descendants (nodes and/or tips). If \code{node.pos = 2}, then the
ordinate of a node is the mean of the ordinates of all the tips of
which it is the ancestor. If \code{node.pos = NULL} (the default),
then its value is determined with respect to other options: if
\code{type = "phylogram"} then `node.pos = 1'; if \code{type =
"cladogram"} and \code{use.edge.length = FALSE} then `node.pos = 2';
if \code{type = "cladogram"} and \code{use.edge.length = TRUE} then
`node.pos = 1'. Remember that in this last situation, the branch
lengths make sense when projected on the x-axis.
If \code{adj} is not specified, then the value is determined with
respect to \code{direction}: if \code{direction = "leftwards"} then
\code{adj = 1} (0 otherwise).
If the arguments \code{x.lim} and \code{y.lim} are not specified by the
user, they are determined roughly by the function. This may not always
give a nice result: the user may check these values with the
(invisibly) returned list (see ``Value:'').
If you use \code{align.tip.label = TRUE} with \code{type = "fan"}, you
will have certainly to set \code{x.lim} and \code{y.lim} manually.
If you resize manually the graphical device (windows or X11) you may
need to replot the tree.
}
\note{
The argument \code{asp} cannot be passed with \code{\dots}.
}
\value{
\code{plot.phylo} returns invisibly a list with the following
components which values are those used for the current plot:
\item{type}{}
\item{use.edge.length}{}
\item{node.pos}{}
\item{node.depth}{}
\item{show.tip.label}{}
\item{show.node.label}{}
\item{font}{}
\item{cex}{}
\item{adj}{}
\item{srt}{}
\item{no.margin}{}
\item{label.offset}{}
\item{x.lim}{}
\item{y.lim}{}
\item{direction}{}
\item{tip.color}{}
\item{Ntip}{}
\item{Nnode}{}
\item{root.time}{}
\item{align.tip.label}{}
}
\author{Emmanuel Paradis, Martin Smith, Damien de Vienne}
\references{
van der Ploeg, A. (2014) Drawing non-layered tidy trees in linear
time. \emph{Journal of Software: Practice and Experience}, \bold{44},
1467--1484.
}
\seealso{
\code{\link{read.tree}}, \code{\link{trex}}, \code{\link{kronoviz}},
\code{\link{add.scale.bar}}, \code{\link{axisPhylo}},
\code{\link{nodelabels}}, \code{\link{edges}},
\code{\link[graphics]{plot}} for the basic plotting function in R
}
\examples{
### An extract from Sibley and Ahlquist (1990)
cat("(((Strix_aluco:4.2,Asio_otus:4.2):3.1,",
"Athene_noctua:7.3):6.3,Tyto_alba:13.5);",
file = "ex.tre", sep = "\n")
tree.owls <- read.tree("ex.tre")
plot(tree.owls)
unlink("ex.tre") # delete the file "ex.tre"
### Show the types of trees.
layout(matrix(1:6, 3, 2))
plot(tree.owls, main = "With branch lengths")
plot(tree.owls, type = "c")
plot(tree.owls, type = "u")
plot(tree.owls, use.edge.length = FALSE, main = "Without branch lengths")
plot(tree.owls, type = "c", use.edge.length = FALSE)
plot(tree.owls, type = "u", use.edge.length = FALSE)
layout(1)
data(bird.orders)
### using random colours and thickness
plot(bird.orders,
edge.color = sample(colors(), length(bird.orders$edge)/2),
edge.width = sample(1:10, length(bird.orders$edge)/2, replace = TRUE))
title("Random colours and branch thickness")
### rainbow colouring...
X <- c("red", "orange", "yellow", "green", "blue", "purple")
plot(bird.orders,
edge.color = sample(X, length(bird.orders$edge)/2, replace = TRUE),
edge.width = sample(1:10, length(bird.orders$edge)/2, replace = TRUE))
title("Rainbow colouring")
plot(bird.orders, type = "c", use.edge.length = FALSE,
edge.color = sample(X, length(bird.orders$edge)/2, replace = TRUE),
edge.width = rep(5, length(bird.orders$edge)/2))
segments(rep(0, 6), 6.5:1.5, rep(2, 6), 6.5:1.5, lwd = 5, col = X)
text(rep(2.5, 6), 6.5:1.5, paste(X, "..."), adj = 0)
title("Character mapping...")
plot(bird.orders, "u", font = 1, cex = 0.75)
data(bird.families)
plot(bird.families, "u", lab4ut = "axial", font = 1, cex = 0.5)
plot(bird.families, "r", font = 1, cex = 0.5)
### cladogram with oblique tip labels
plot(bird.orders, "c", FALSE, direction = "u", srt = -40, x.lim = 25.5)
### facing trees with different informations...
tr <- bird.orders
tr$tip.label <- rep("", 23)
layout(matrix(1:2, 1, 2), c(5, 4))
plot(bird.orders, "c", FALSE, adj = 0.5, no.margin = TRUE, label.offset = 0.8,
edge.color = sample(X, length(bird.orders$edge)/2, replace = TRUE),
edge.width = rep(5, length(bird.orders$edge)/2))
text(7.5, 23, "Facing trees with\ndifferent informations", font = 2)
plot(tr, "p", direction = "l", no.margin = TRUE,
edge.width = sample(1:10, length(bird.orders$edge)/2, replace = TRUE))
### Recycling of arguments gives a lot of possibilities
### for tip labels:
plot(bird.orders, tip.col = c(rep("red", 5), rep("blue", 18)),
font = c(rep(3, 5), rep(2, 17), 1))
plot(bird.orders, tip.col = c("blue", "green"),
cex = 23:1/23 + .3, font = 1:3)
co <- c(rep("blue", 9), rep("green", 35))
plot(bird.orders, "f", edge.col = co)
plot(bird.orders, edge.col = co)
layout(1)
## tidy trees
tr <- rtree(100)
layout(matrix(1:2, 2))
plot(tr)
axis(2)
plot(tr, "t")
axis(2)
## around 20 percent gain on the y-axis
}
\keyword{hplot}
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