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## drop.tip.R (2023-01-09)
## Remove Tips in a Phylogenetic Tree
## Copyright 2003-2023 Emmanuel Paradis, 2017-2023 Klaus Schliep, 2018 Joseph Brown
## This file is part of the R-package `ape'.
## See the file ../COPYING for licensing issues.
keep.tip <- function(phy, tip, ...) UseMethod("keep.tip")
keep.tip.phylo <- function(phy, tip, ...)
{
Ntip <- length(phy$tip.label)
## convert to indices if strings passed in
if (is.character(tip)) {
idx <- match(tip, phy$tip.label)
## stop on bad tip names
## alternative is to warn but proceed. not sure what stance is
if (anyNA(idx)) {
um <- c("unmatched tip labels:\n", paste(tip[is.na(idx)], collapse = " "))
stop(um)
}
tip <- idx
} else {
# check that passed in indices are all valid
out.of.range <- tip > Ntip
if (any(out.of.range)) {
warning("some tip numbers were larger than the number of tips: they were ignored")
tip <- tip[!out.of.range]
}
}
## get complement tip indices to drop
toDrop <- setdiff(1:Ntip, tip)
drop.tip(phy, toDrop, ...)
}
extract.clade <- function(phy, node, root.edge = 0, collapse.singles = TRUE, interactive = FALSE)
{
n <- length(phy$tip.label)
if (interactive) {
cat("Click close to the node...\n")
node <- identify(phy)$nodes
} else {
if (length(node) > 1) {
node <- node[1]
warning("only the first value of 'node' has been considered")
}
if (is.character(node)) {
if (is.null(phy$node.label))
stop("the tree has no node labels")
node <- match(node, phy$node.label) + n
if (is.na(node)) stop("'node' not among the node labels.")
}
if (node <= n)
stop("node number must be greater than the number of tips")
}
if (node == n + 1L) return(phy)
keep <- prop.part(phy)[[node - n]]
drop.tip(phy, (1:n)[-keep], root.edge = root.edge, rooted = TRUE,
collapse.singles = collapse.singles)
}
drop.tip <- function(phy, tip, ...) UseMethod("drop.tip")
drop.tip.phylo <- function(phy, tip, trim.internal = TRUE, subtree = FALSE,
root.edge = 0, rooted = is.rooted(phy), collapse.singles = TRUE,
interactive = FALSE, ...)
{
Ntip <- length(phy$tip.label)
## find the tips to drop:
if (interactive) {
cat("Left-click close to the tips you want to drop; right-click when finished...\n")
xy <- locator()
nToDrop <- length(xy$x)
tip <- integer(nToDrop)
lastPP <- get("last_plot.phylo", envir = .PlotPhyloEnv)
for (i in 1:nToDrop) {
d <- sqrt((xy$x[i] - lastPP$xx)^2 + (xy$y[i] - lastPP$yy)^2)
tip[i] <- which.min(d)
}
} else {
if (is.character(tip))
tip <- which(phy$tip.label %in% tip)
}
out.of.range <- tip > Ntip
if (any(out.of.range)) {
warning("some tip numbers were larger than the number of tips: they were ignored")
tip <- tip[!out.of.range]
}
if (!length(tip)) return(phy)
if (length(tip) == Ntip) {
if (Nnode(phy) < 3 || trim.internal) { # by Klaus (2018-06-21)
warning("drop all tips of the tree: returning NULL")
return(NULL)
}
}
wbl <- !is.null(phy$edge.length)
if (length(tip) == Ntip - 1 && trim.internal) {
i <- which(phy$edge[, 2] == (1:Ntip)[-tip])
res <- list(edge = matrix(2:1, 1, 2),
tip.label = phy$tip.label[phy$edge[i, 2]],
Nnode = 1L)
class(res) <- "phylo"
if (wbl) res$edge.length <- phy$edge.length[i]
if (!is.null(phy$node.label))
res$node.label <- phy$node.label[phy$edge[i, 1] - Ntip]
return(res)
}
if (!rooted) {
phy$root.edge <- NULL # moved from below (2021-09-29)
if (subtree) {
phy <- root(phy, (1:Ntip)[-tip][1])
root.edge <- 0
}
}
phy <- reorder(phy)
NEWROOT <- ROOT <- Ntip + 1
Nnode <- phy$Nnode
Nedge <- dim(phy$edge)[1]
if (subtree) {
trim.internal <- TRUE
tr <- reorder(phy, "postorder")
N <- .C(node_depth, as.integer(Ntip),
as.integer(tr$edge[, 1]), as.integer(tr$edge[, 2]),
as.integer(Nedge), double(Ntip + Nnode), 1L)[[5]]
}
edge1 <- phy$edge[, 1] # local copies
edge2 <- phy$edge[, 2] #
keep <- !logical(Nedge)
## delete the terminal edges given by `tip':
keep[match(tip, edge2)] <- FALSE
if (trim.internal) {
ints <- edge2 > Ntip
## delete the internal edges that do not have anymore
## descendants (ie, they are in the 2nd col of `edge' but
## not in the 1st one)
repeat {
sel <- !(edge2 %in% edge1[keep]) & ints & keep
if (!sum(sel)) break
keep[sel] <- FALSE
}
if (subtree) {
## keep the subtending edge(s):
subt <- edge1 %in% edge1[keep] & edge1 %in% edge1[!keep]
keep[subt] <- TRUE
}
if (root.edge && wbl) {
degree <- tabulate(edge1[keep])
if (degree[ROOT] == 1) {
j <- integer(0) # will store the indices of the edges below the new root
repeat {
i <- which(edge1 == NEWROOT & keep)
j <- c(i, j)
NEWROOT <- edge2[i]
## degree <- tabulate(edge1[keep]) # utile ?
if (degree[NEWROOT] > 1) break
}
keep[j] <- FALSE
## if (length(j) > root.edge) j <- 1:root.edge
j <- j[1:root.edge]
NewRootEdge <- sum(phy$edge.length[j])
if (length(j) < root.edge && !is.null(phy$root.edge))
NewRootEdge <- NewRootEdge + phy$root.edge
phy$root.edge <- NewRootEdge
}
}
}
##if (!root.edge) phy$root.edge <- NULL # moved above (2021-09-29)
## drop the edges
phy$edge <- phy$edge[keep, ]
if (wbl) phy$edge.length <- phy$edge.length[keep]
## find the new terminal edges (works whatever 'subtree' and 'trim.internal'):
TERMS <- !(phy$edge[, 2] %in% phy$edge[, 1])
## get the old No. of the nodes and tips that become tips:
oldNo.ofNewTips <- phy$edge[TERMS, 2]
## in case some tips are dropped but kept because of 'subtree = TRUE':
if (subtree) {
i <- which(tip %in% oldNo.ofNewTips)
if (length(i)) {
phy$tip.label[tip[i]] <- "[1_tip]"
tip <- tip[-i]
}
}
n <- length(oldNo.ofNewTips) # the new number of tips in the tree
## the tips may not be sorted in increasing order in the
## 2nd col of edge, so no need to reorder $tip.label
phy$edge[TERMS, 2] <- rank(phy$edge[TERMS, 2])
## fix by Thomas Sibley (2017-10-28):
if (length(tip)) phy$tip.label <- phy$tip.label[-tip]
## make new tip labels if necessary:
if (subtree || !trim.internal) {
## get the numbers of the nodes that become tips:
node2tip <- oldNo.ofNewTips[oldNo.ofNewTips > Ntip]
## fix by Thomas Sibley (2017-10-28):
new.tip.label <-
if (!length(node2tip)) {
character(0)
} else if (subtree && is.null(phy$node.label)) {
paste("[", N[node2tip], "_tips]", sep = "")
} else {
if (is.null(phy$node.label)) rep("NA", length(node2tip))
else phy$node.label[node2tip - Ntip]
}
# if (!is.null(phy$node.label))
# phy$node.label <- phy$node.label[-(node2tip - Ntip)]
phy$tip.label <- c(phy$tip.label, new.tip.label)
}
phy$Nnode <- dim(phy$edge)[1] - n + 1L # update phy$Nnode
## The block below renumbers the nodes so that they conform
## to the "phylo" format
newNb <- integer(Ntip + Nnode)
newNb[NEWROOT] <- n + 1L
sndcol <- phy$edge[, 2] > n
newNb[sort(phy$edge[sndcol, 2])] <- (n + 2):(n + phy$Nnode)
phy$edge[sndcol, 2] <- newNb[phy$edge[sndcol, 2]]
phy$edge[, 1] <- newNb[phy$edge[, 1]]
storage.mode(phy$edge) <- "integer"
if (!is.null(phy$node.label)) # update node.label if needed
phy$node.label <- phy$node.label[which(newNb > 0) - Ntip]
if (collapse.singles) phy <- collapse.singles(phy)
phy
}
## "multiPhylo" methods by Klaus:
keep.tip.multiPhylo <- function(phy, tip, ...)
{
if (is.null(attr(phy, "TipLabel"))) {
tmp <- try(.compressTipLabel(phy), TRUE)
if (!inherits(tmp, "try-error")) phy <- tmp
}
if (!is.null(attr(phy, "TipLabel"))) {
phy <- lapply(phy, keep.tip, tip, ...)
class(phy) <- "multiPhylo"
phy <- .compressTipLabel(phy)
} else {
if (!inherits(tip, "character"))
stop("Trees have different labels, tip needs to be of class character!")
phy <- lapply(phy, keep.tip, tip, ...)
class(phy) <- "multiPhylo"
}
phy
}
drop.tip.multiPhylo <- function(phy, tip, ...)
{
interactive <- if (hasArg(interactive)) list(...)$interactive else FALSE
if (interactive)
stop("interactive = TRUE does not work for drop.tip.multiPhylo().")
if (is.null(attr(phy, "TipLabel"))) {
tmp <- try(.compressTipLabel(phy), TRUE)
if (!inherits(tmp, "try-error")) phy <- tmp
}
if (!is.null(attr(phy, "TipLabel"))) {
phy <- lapply(phy, drop.tip, tip, ...)
class(phy) <- "multiPhylo"
phy <- .compressTipLabel(phy)
} else {
if (!inherits(tip, "character"))
stop("Trees have different labels, tip needs to be of class character!")
phy <- lapply(phy, drop.tip, tip, ...)
class(phy) <- "multiPhylo"
}
phy
}
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